TYPE: Costa Rica: [Alajuela:] Bois humides à San Pedro de San Ramón, 1150 m, Jul. 1921, A.M. Brenes 76 (holotype, B, destroyed; drawings of the holotype, AMES; detail of the drawing, floral dissections, AMES; detail of the drawing, habit, AMES. Neotype, designated by Barringer, 1984: Costa Rica. Alajuela: San Pedro de San Ramon, 1100 m, Sept. 1921, A.M. Brenes 121, AMES; isoneotype, CR).